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Derrick

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  1. Further to this thread. Here are two herbarium specimens attributed to H. angustifolium by Adolph D. E. Elmer in 1910. Collection not type. Elmer (1910) Philippines, Sibuyan Island, Capiz Province, Romblon, Magallanes (Mt. Giting-Giting) http://data.biodiversitydata.nl/naturalis/specimen/U.1561946 http://data.biodiversitydata.nl/naturalis/specimen/L.2913092 Update. Jebb & C. R. Huxley have referred these two specimens to H. formicarum in their recent revision. I am not convinced.
  2. Note these four names are listed as new synonyms on page 41. Hydnophytum kochii Val. (1911) syn. nov. H. ledermannii Val. (1927) syn. nov. H. nigrescens Merr. & L.M.Perry (1945) syn. nov. H. contortum Merr. & L.M.Perry (1945) syn. nov. Three of these are also listed as var. nov. H. ledermannii p44, H. nigrescens p45, and H. contortum p43. H. kochii is shown as a synonym of var. petiolatum on p42 but is not a new variety. I presume this is correct nomenclature, but it is confusing.
  3. H. virgatum Val. (Theodoric Valeton) Just's botanischer jahresbericht. Systematisch geordnetes repertorium der botanischen literatur aller länder, vol.61, p142, (1927). (Bot. Jahrb. Syst.), not yet digitised. Update. Jebb & C. R. Huxley, The tuberous epiphytes of the Rubiaceae 7: a revision of the genus Hydnophytum, Blumea 64, p86, (2019.) With illustration and a member of their Uncertain and Little-Known Species. Type, Ledermann 11646, 11701, 11849 (Bogor Herbarium not seen, presumed lost), Papua New Guinea, East Sepik Province, Schrader Mountains. I have found a more recent collection claimed to be this species using Valeton’s key, but these are not types. Takeuchi (1995) Papua New Guinea, Madang Province, Bismarck Range. http://data.biodiversitydata.nl/naturalis/specimen/L%20%200074528 And. http://data.biodiversitydata.nl/naturalis/specimen/L.2913882 Collection notes. Ant-house epiphyte, tuber brown, completely obscured by roots and debris, leaves coriaceous, dark green, mid green below, inflorescence axillary, sessile, pauciflorous, flower 4 merous, yellow. Papua New Guinea, Madang Province, ridge overlooking campsite 4, at 600-900 m.
  4. H. subsessile Val. (Theodoric Valeton) Just's botanischer jahresbericht. Systematisch geordnetes repertorium der botanischen literatur aller länder, vol.61, p142, (1927). (Bot. Jahrb. Syst.), not yet digitised. Update. Jebb & C. R. Huxley, in The tuberous epiphytes of the Rubiaceae 7: a revision of the genus Hydnophytum, Blumea 64, p86, (2019.) A member of their Uncertain and Little Known Species. Type, Lauterbach 3164, (B not seen, presumed lost), Papua New Guinea. The only herbarium placement I can find is an endorsement of H. linearifolium var. subsessile on the type sheet for H. linearifolium. I repeat here. Type, Ledermann 9700. (1912) appears to have been labelled H. linearifolium var. subsessile but the writing is not clear. http://data.biodiversitydata.nl/naturalis/specimen/L%20%200000491 And https://plants.jstor.org/stable/10.5555/al.ap.specimen.l0000491
  5. H. stenophyllum Val. (Theodoric Valeton) Just's botanischer jahresbericht. Systematisch geordnetes repertorium der botanischen literatur aller länder, vol.61, p140, (1927). (Bot. Jahrb. Syst.), not yet digitised. Update. Jebb & C. R. Huxley, in The tuberous epiphytes of the Rubiaceae 7: a revision of the genus Hydnophytum, Blumea 64, p85, (2019.) A member of their Uncertain and Little Known Species. Type, Schlechter 18173 (1908) Jebb & C. R. Huxley note “K not seen, presumed lost” Papua New Guinea, Madang Province, Huon Peninsula, Finisterre Mountains at 300m (984 ft.) Yet here it is. http://apps.kew.org/herbcat/getImage.do?imageBarcode=K000761951 This specimen has distinct, long and somewhat narrow leaves. Merr. & L.M. Perry note when referring to H. ramispinum. “In leaf-size and flower, this species (i.e. ramispinum) is near Hydnophytum stenophyllum Val. and H. punamense Lauterb. In both the latter species the leaves are either attenuate-acute or acute, and in neither is there any mention of the furfuraceous character of the young bark. Unfortunately, many of the species have no indication of the characters of the tuberous base. (Elmer Drew Merrill & Lily May Perry) Journal Arnold Arboretum Vol 26, pp23, (1945) (J. Arnold Arbor.) http://www.biodiversitylibrary.org/page/8443412#page/28/mode/1up
  6. Jeff. That drawing of a specimen in FLOWER represents a TYPE of the basionym. The following sentence is from my opening post of this thread. "Carnevali (2009) also states that: "This beautiful species was described by James Bateman in his classic book 'The Orchidaceae of Mexico and Guatemala' published as pl.30, in 1843 as Schomburgkia tibicinis, from plants collected by George Ure Skinner in Guatemala. This is correct," This URL links to THE holotype of the basionym Schomburgkia tibicinis with a description and further notes in Latin and English. Thus, even if you wish to ignore the expert work of Rolfe and use the basionym, the following flower is correct for tibicinis and Robert's flower is not. "Petals nearly equal, very much curled, upwards of an inch long, dark chestnut brown inside, and dirty purple without." http://www.biodiversitylibrary.org/item/15471#page/105/mode/1up
  7. Hello Andreas. The links normally work but their server seems to be having problems. Try this method. but it too is currently having problems. Click on www.philippine plants.org Then click on Angiosperm Families. Scroll to Rubiaceae, then to Hydnophytum. I see that Dr Pieter Pelser has already ascribed most names to H. formicarum or H. moseleyanum after Jebb & Huxley 2019. H. angustifolium is kept separate.
  8. A while back I suggested to Co's Digital Flora of the Philippines that some photos in their database may represent this species. Jebb & C. R. Huxley (2019) have it listed under their little known species. I suspect this may be a rare situation where we amateurs can provide professionals with new evidence. https://www.facebook.com/groups/philippineplants/ Incidentally, this site is run from little New Zealand. H. angustifolium Merr. (Elmer Drew Merrill) The Philippine Journal of Science. Section C, Botany 3, p162, (1908) (Philipp. J. Sci., C.) Descriptions in Latin as then essential, and in English. http://www.biodiversitylibrary.org/page/673338#page/185/mode/1up Merr. Leaflets Philippine Botany, Vol 3, Art 54, p1038, in English. (Leafl. Philipp. Bot.) http://www.biodiversitylibrary.org/page/776329#page/277/mode/1up Merr. Notes. A very characteristic species, readily recognizable by its narrow sessile leaves, the nerves of which are obsolete or nearly so, and its elongated fruit. Tuber irregular, at least 15 cm (6 inches) in diameter, brown or grayish, unarmed. Stems several, diffusely branched, at least 60 cm long, gray or brown, slender, the branches elongated, younger ones, brown, furfuraceous (covered with flaky scales), somewhat angled. Leaves lanceolate or narrowly lanceolate, 5-10 cm (2-4 inches) long when dry, somewhat shining, gradually narrowed at base and apex, tip acute or blunt, midrib prominent beneath, lateral nerves obsolete or very obscure. Flowers fascicled (bundled) in axils, white. Calyx cylindrical 1 mm., long and wide, truncate. Corolla 2 mm long, inside slightly barbate at the middle. Anthers 0.7 mm long. Style 1-2 mm long. Fruit red, somewhat fleshy when fresh, to 1 cm long, about 4 mm m diameter at the base, gradually narrowed upward, apparently 1-celled with a single seed. Update. Jebb & C. R. Huxley (2019) were unable to place this in their concepts of H. moseleyanum or H. formicarum or “most especially H. puffii” due to a lack of herbarium specimens, that are possibly lost. Habitat/Range. A forest epiphyte at 200-600 m (656-1968.5 ft.), Mostly in the southern islands, yet common on central Sibuyan Island. Zamboanga & Lanao Del Sur Provinces, Mindanao Island, Philippines. The following images possibly represent this species. http://131.230.176.4/imgs/pelserpb/r/Rubiaceae_Hydnophytum_plant2_49811.html http://131.230.176.4/imgs/pelserpb/r/Rubiaceae_Hydnophytum_plant2_49820.html http://131.230.176.4/imgs/pelserpb/r/Rubiaceae_Hydnophytum_plant2_49818.html http://131.230.176.4/imgs/pelserpb/r/Rubiaceae_Hydnophytum_plant2_49817.html
  9. This is an authentic type image of Myrmecophila tibicinis. The above image is not tibicinis under whatever historic genus one wishes to place it. http://www.botanicus.org/page/769219 This type image was referenced in my opening post but the URL used then no longer works.
  10. As I update Epiphytic Myrmecophytes: Bizarre Wonder of Nature 2019 (currently almost through chapter 9, Hydnophytum, I uncovered quite a number of hints of what to expect in a revision. About a week ago someone on the WWW reported that this revision was imminent but I found that vary hard to believe. We had waited soooo long. I now have to re-evaluate this entire chapter that has already involved many months of work. It has been designed primarily as a searchable a database, supported with photographs wherever possible. So it probably still has value for a wide range of enthusiasts. I do wonder how much will change as botanists apply ever more molecular genetics in their work; for example, some of what are only suggested manuscript names may need to be resurrected. My database probably has most of them.
  11. The biodiversity. naturalis site is closed for some time due to reorganization. However, access to Hydnophytum sheets can be achieved through https://www.europeana.eu/portal/en/#
  12. H. radicans Becc. (Odoardo Beccari) in, Malesia raccolta 2, p132, (1885.) Herbarium specimens assigned to this name show large variations in leaf shapes. Do we have cryptic species here? (Malesia 2) http://www.biodiversitylibrary.org/page/44197640#page/246/mode/1up In summary http://www.biodiversitylibrary.org/page/44197640#page/237/mode/1up Beccari’s image http://www.biodiversitylibrary.org/page/44197829#page/147/mode/1up Beccari illustration http://plantillustrations.org/illustration.php?id_illustration=327448 Val. (Theodoric Valeton) in Nova Guinea: résultats de l'expédition scientifique néerlandaise à la Nouvelle-Guinée. (Nova Guinea 8, part1), p503. (1906-1907.) http://www.biodiversitylibrary.org/item/42443#page/535/mode/1up Merr. & L. M. Perry. (Elmer Drew Merrill & Lily May Perry) Journal of the Arnold Arboretum vol. 26, p24, (1945). (J. Arnold Arbor.) It may not always form a tuber, See page 17 under H. archboldiana in, https://www.biodiversitylibrary.org/page/8443412#page/28/mode/1up A common canopy epiphyte in ridge forest, tuber small, branches long, weak, semi-herbaceous, leaves fleshy. Flowers white, (slightly yellow to ochre according to Beccari, fruit soft, red, ± 5 mm. long, 3 mm. Ø. Collections. Papua New Guinea, Northern Province, Buna District. Western Province, Palmer River, 2 miles below junction of Black River, at 100 m. (328 ft.) http://www.pngplants.org/ Papua New Guinea, East Sepik Province, Andai. Type, Florence, H. http://plants.jstor.org/stable/10.5555/al.ap.specimen.fi008893 (Undated) Versteeg collections, Nova Guinea, neerlandica meridionalis. http://data.biodiversitydata.nl/naturalis/specimen/L.2913925 http://data.biodiversitydata.nl/naturalis/specimen/L.2913926 (1926) Docters van Leeuwen coll. West New Guinea Island, Nassau Mts. Taritaku (Rouffaer) River forest. http://data.biodiversitydata.nl/naturalis/specimen/L.2913974 (1926) Docters van Leeuwen coll. West New Guinea Island, Albatros bivak (camp) Mamberamo River. Flower yellow. http://data.biodiversitydata.nl/naturalis/specimen/L.2913949 (1935 Carr., coll.) Papua New Guinea, Central Province, Rouna. Rouna falls on the Laloki river are a popular tourist site on the road up to the Sogeri Plateau behind Port Moresby. http://data.biodiversitydata.nl/naturalis/specimen/L.2913951 http://data.biodiversitydata.nl/naturalis/specimen/L.2913916 (1936 Carr., coll.) Papua New Guinea, Oro Province, Kokoda. http://data.biodiversitydata.nl/naturalis/specimen/L.2913971 (1956) Papua New Guinea, Milne Bay Province, Normanby Island, Lebudowa River. http://data.biodiversitydata.nl/naturalis/specimen/L.2913953 (1963) Epiphyte in branches of Samanea saman, large tuberous base, sparsely galleried, foliage bearing shoots up to 7ft long, slender but erect. Flowers white, fruit orange-red when ripe. Papua New Guinea, Morobe Province, Morobe Agricultural station. 3 miles from LAE. http://apps.kew.org/herbcat/detailsQuery.do?barcode=K001045937 (1963) Shrub, 3 ft., pendulous, bulbous base housing ants. Flower white. Fruit red, succulent. Papua New Guinea, Morobe Province, Bumbu River. http://data.biodiversitydata.nl/naturalis/specimen/L.2914127 (1975) Epiphyte, swollen chambered stem, stems dark brown, leaves leathery, dark green above, paler beneath, inflorescence, pale grey-green, flower buds white; young fruits orange. At 250 m in lowland rainforest of Terminalia, Pometia, Elaeocarpus & Albizia. Kikori District, Gulf Province; 200 k. from coast on Purari river; south side of ridge upstream from Wabo Creek. http://apps.kew.org/herbcat/detailsQuery.do?barcode=K001045938 (1975) C. R. Huxley & Kirk coll. Tuber 53x23 cm., brown, smooth with longitudinal ridges, Leaves mid green, Flowers white, fruit red. Papua New Guinea, Central Province, Varirata National Park. http://data.biodiversitydata.nl/naturalis/specimen/L.2913948 (1994) Indonesia, Maluku Province, (Moluccas Islands) Aru Archipelago (the most eastern group close to West New Guinea Island) Kobroor Island, 8 km SE of Jirlay. http://data.biodiversitydata.nl/naturalis/specimen/L.2914173 (1995) Roots tuberous, ant-inhabited, leaves thin, leathery, fruit red. In secondary forest on limestone hill at 500 m. (1640 ft.) West New Guinea Island, West Papua Province, Bird’s Head Peninsula, surroundings of Ayawasi. http://data.biodiversitydata.nl/naturalis/specimen/L.2914138 (2000) Ant home slightly warty and bumpy but relatively smooth with no ridges or spines; smelling of a potato when cut; bulbous; to 9 x 5.5 cm; with mass of fibrous roots around the base; several young plants germinating in root mass, young plants already with small hollow ant houses and ants; ants not biting, small and black. Stems pendulous, very pale grey, rooting sparsely along their length especially at base; main stem 63 cm long; secondary stem a branch 36 cm long arising 11 cm along the main stem; stems ridged, ridge arising from axil of a lower node moving around stem and ending at stipule (or vice versa). Leaves orientated in horizontal pale, i.e. petioles twisting lime green above, rusty orange green below. Inflorescence a dichasial cyme branching 3 times, i.e. with 8 terminal inflorescence heads (all inflorescences branched exactly the same and at the same stage - anthesis nearly finished, mostly all young fruits, only one mature fruit seen); inflorescence stalk woody. Corolla translucent white, tinged pale orange at apex (RHS colour chart 25C), corolla appendages very tufted, pure white. Young fruits orange-green, orange at apex (RHS 24A); ripe fruit orange-red (RHS 34A). West New Guinea, Papua Province, Mimika Regency. Kuala Kencana, in front of Rima Irian Golf Club towards access road. http://data.biodiversitydata.nl/naturalis/specimen/L.2914122 Specimens originally named as, H. “communis” (1920 Dr. Lam collection, West New Guinea, Mamberamo River, Pionierbivak. http://data.biodiversitydata.nl/naturalis/specimen/L.2913968 http://data.biodiversitydata.nl/naturalis/specimen/L.2913928 http://data.biodiversitydata.nl/naturalis/specimen/L.2913933 H. “communis” (1920 Lam coll.) Mamberamo River, pr. Meervlakte bij Prauwenbivak. http://data.biodiversitydata.nl/naturalis/specimen/L.2913929 See also H. radicans var communis below. H. kelelense Val. Not type but may become a synonym, Papua New Guinea, Morobe Province, Along the Bunga River about 15 miles NE of Lae. (Hartley coll. 1963.) http://data.biodiversitydata.nl/naturalis/specimen/L.2913931 H. montis-kani Val. (1953 Brass coll.) Papua New Guinea; Milne Bay District. Biniguni Camp, Gwariu River. Low epiphyte in Riverine rain forest at 200 m. (656 ft.) (See (Brass 1953) above. http://data.biodiversitydata.nl/naturalis/specimen/L.2913958 H. montis-kani Val. (1972 Katik coll.) Lae - Leiden - Canberra Botanical Expedition. Papua New Guinea, Milne Bay Province, Rabaraba, Ma-u River, Mt Suckling Complex. http://data.biodiversitydata.nl/naturalis/specimen/L.2913939 H. normale Becc. (1935 Clemens coll. not type) Papua New Guinea, Morobe Province, Sattelberg. http://data.biodiversitydata.nl/naturalis/specimen/L.2913970 H. radicans Becc var “communis” Val. (1920, Lam coll.) West New Guinea Island, Papua Province, Mamberamo River, at 70 m. (230 ft.) Pionierbivak (one of a number of camps along explored rivers.) See bottom of p104, Biogeography and Ecology of New Guinea. Which details early explorations of New Guinea and is digitised. http://data.biodiversitydata.nl/naturalis/specimen/L.2913934 H. radicans Becc var communis Val. (1912 Pulle collection) West New Guinea Island, Papua Province (south), Lorentz River, Kloofbivak (Kloof Camp) http://data.biodiversitydata.nl/naturalis/specimen/L.2913967 H. “thomsonii” (1914 Thompson coll.) West New Guinea, Nova Guinea neerlandica in reg. flum. Mamberamo, pr. flur., Utkenrivier op Kleischiefer. http://data.biodiversitydata.nl/naturalis/specimen/L.2913938 Probable synonyms. H. amplifolium S. Moore, Isotype (1926) published in Journal of Botany British & Foreign LXV (65) p270, (1927.) http://plants.jstor.org/stable/10.5555/al.ap.specimen.bri-aq0570115 H. dolichophyllum Val. (1912, Ledermann collection, not type. Huxley refers to H. radicans. Papua New Guinea, Sepikgebiet. http://data.biodiversitydata.nl/naturalis/specimen/L.2913930 H. keiense Becc., (1873) Type, Moluccas Island, Kai (Kei) Island, kei bandan a weri. http://apps.kew.org/herbcat/getImage.do?imageBarcode=K000761921 H. keiense Becc. Papua New Guinea, Milne Bay Province, Peria Creek, Kwagira (Kwaria) River 9°42'S 149°23'E. Brass collection (1953). http://kiki.huh.harvard.edu/databases/specimen_search.php?mode=details&id=518743 H. punamense Lauterb. (Carr 1936) Flowers white, fruit dull orange, Papua New Guinea, Central Province, Isuarava, 4 20 S, 152 16 E, http://apps.kew.org/herbcat/detailsQuery.do?barcode=K001045863 And. http://data.biodiversitydata.nl/naturalis/specimen/L.2913932 http://data.biodiversitydata.nl/naturalis/specimen/L.2913924 H. punamense Lauterb. (Brass 1953) High epiphyte, branches 60-70 cm long, sub-pendent from unarmed tuberous base ± 20 cm in diameter, flowers white, fruit orange. Peria Creek, Kwagira River, Milne Bay District, Papua New Guinea. http://apps.kew.org/herbcat/detailsQuery.do?barcode=K001045936 H. punamense Lauterb. (1964) leaves mid green torus (tuber?) green, flowers white, epiphyte at 4 ft. on Sonneratia overhanging fresh water lagoon, Papua New Guinea. Sepik District, Leitre village. http://apps.kew.org/herbcat/detailsQuery.do?barcode=K001045862 A very suspect identification. (1968 coll.) Borneo Island, Malaysia, Sabah State, Madai caves, Madai Baturong Forest Reserve at 100 m. (328 ft.). http://data.biodiversitydata.nl/naturalis/specimen/L.2913875
  13. H. dolichophyllum Val. (Theodoric Valeton) in Just’s botanischer jahresbericht. Systematisch geordnetes repertorium der botanischen literatur aller länder, vol.61, p130, (1927) (Bot. Jahrb. Syst.) Only digitised to 1922. Here an H. dolichophyllum Val., collection (not type, probably because they were lost during the second world war) is referred to H. radicans by Huxley in 1991. (Ledermann collection 1912), Papua New Guinea, Sepikgebiet.
  14. Jeff. I suggest you check a lot more herbarium sheets. I have already checked at least 100 sheets of radicans and possibly associated species. I found that the leaf shape of species accepted as H. radicans by the experts is surprisingly LARGE. To the degree that leaf shape has very little influence on the defining features of this species. My initial thought was that DNA evidence might change our understanding of what defines H. radicans. Yet Dr Guillaume Chomicki's acceptance of the H. kiense specimen above, suggests NO. He certainly is using DNA in his work, but I cannot guarantee DNA tests were performed on this particular specimen.
  15. (1968 coll.) Borneo Island, Malaysia, Sabah State, Madai caves, Madai Baturong Forest Reserve at 100 m. (328 ft.). http://data.biodiversitydata.nl/naturalis/specimen/L.2913875
  16. Here a Kew Type specimen originally labelled H. keiense is assigned to H. radicans by Huxley 1992. Weri Bandan, Kai Besar (formerly Kei) Island, south east Moluccas archipelago (Spice Islands), Indonesia, (Isole Kei, Bandan a Weri. Isole = island, Kai Besar being the largest. Bandan is a political/population centre, Weri means stream. Note the low number of collections ever assigned to keiense or kejense. http://apps.kew.org/herbcat/getImage.do?imageBarcode=K000761921 H. keiense Becc. Papua New Guinea, Milne Bay Province, Peria Creek, Kwagira (Kwaria) River 9°42'S 149°23'E. Brass collection (1953). Here assigned to H. radicans by Chomicki in 2013. http://kiki.huh.harvard.edu/databases/specimen_search.php?mode=details&id=518743 Thus it seems very probable that the names keiense and kejense will become historical when a hydnophytum revision is finally published.
  17. H. ovatum Miq. (Friedrich Anton Wilhelm Miquel) in Annales Musei botanici lugduno-batavi. vol. 4, Select Rubiaceae of The East Indian Archipelago, p257/8, (1869.) (Ann. Mus. Bot. Lugduno-Batavum) http://www.biodiversitylibrary.org/item/2547#page/286/mode/1up Beccari Illustration http://plantillustrations.org/illustration.php?id_illustration=327540. Scroll down to best image. Description. Translated from Latin. “(New) sp. Trunk globose, branches long, alternating and double-branched, acute on the upper surface and obtusely tetragonal on the lower; stipules short, wide, deciduous; leaves sessile or with a very short petiole, rounded oval, wide at the base, slightly notched or truncate, very obtuse at apex or rounded, coriaceous, very smooth, without veins on lower surface, with a pale bloom; flowers axillary, from tubercles. Very distinct species. Trunk globose, 2-inch in diameter, inside structure the same as above, covered with a pitted, wrinkled black cortex, perhaps because of exsiccation (drying). Stems or branches two feet long, or shorter, very ramose and nodose, with blackish wrinkled cortex underneath, dark brown and smooth above, tetragonal with acute angles. Leaves brown-black on the upper surface, with few veins and ribs but distinct, pale greyish white on the bottom surface with no veins or, only rarely, with 2-3 ribs on both sides, 1¼-1¾ inch long. Flowers only incomplete remains are left.” Habitat/Range. Ternate Island, Moluccas Islands (Teysmann & De Vriese collection). http://www.biodiversitylibrary.org/page/42963387#page/567/mode/1up Becc. (Odoardo Beccari) Malesia Raccolta 2, p143, (1884). (Malesia 2.) http://www.biodiversitylibrary.org/page/44197923#page/257/mode/1up Collections. (1859) Isotypes, Teysmann & De Vriese collections, http://data.biodiversitydata.nl/naturalis/specimen/L%20%200057493 http://data.biodiversitydata.nl/naturalis/specimen/L%20%200057494 (1874) Beccari collection http://apps.kew.org/herbcat/getImage.do?imageBarcode=K000761922 (1921) Moluccas Archipelago, N Ternate Island, Foramadiahi Village. http://data.biodiversitydata.nl/naturalis/specimen/L.2913993 (1951) Epiphyte, flower white, in thinned forest, Moluccas Islands, Ternate Island, Puk v. east slope 1500m. (4921 ft.) http://data.biodiversitydata.nl/naturalis/specimen/L.2913996 (1974) Vogel. Gregarious epiphyte, flower white, Open low forest to 20 m (66 ft.) with little undergrowth on hillside of red porous nickel soil. North Moluccas, Obi Isl., W. part, Jikodolong. http://data.biodiversitydata.nl/naturalis/specimen/L.2914166 Undated ?deVriese, Ternate http://data.biodiversitydata.nl/naturalis/specimen/L.2913994 Undated Teysmann, Ternate http://data.biodiversitydata.nl/naturalis/specimen/L.2913995 Undated, Ternate http://data.biodiversitydata.nl/naturalis/specimen/U.1561887
  18. H. alboviride. Merr. & L. M. Perry, (Elmer Drew Merrill & Lily May Perry) in Journal of the Arnold Arboretum, Vol. 26, p21, (1945.) (J. Arnold Arbor.) Type, Brass 12682&3. Merr & Perry note. Of the species of Hydnophytum already described from New Guinea, this one superficially suggests H. Hellwigii (sic) Warb., but differs considerably in particulars. Apart from small leaves, distinctive characters specify, flowers in alveoli surrounded by bracts and long brown hairs; calyx with cilia 0.5 mm. long, also brown; corolla glabrous within; anthers closely affixed by the middle of the back, only apices exserted. On long leafless branches (the leaves often being only toward the tip), flowers or buds with the corolla half protruding may be observed at nodes. http://www.biodiversitylibrary.org/page/8443412#page/25/mode/1up Synonym H. crassicaule, West New Guinea, Mt Dorman (Doormantop) at 3200 m. (10,498 ft.) http://data.biodiversitydata.nl/naturalis/specimen/L.2913818 And. http://data.biodiversitydata.nl/naturalis/specimen/L.2913091 http://data.biodiversitydata.nl/naturalis/specimen/L%20%200000477 Epiphytic on branches of large trees in mossy forest, large plant, branches to 1m. Leaves flat, flowers greenish white, 18 k s. of Bernhard Camp. Duplicate type. http://data.biodiversitydata.nl/naturalis/specimen/L%20%200000474 Originally labelled H. condensatum s. n., then in 1991. H. alboviride, and (undated) holotype of H. vaccinifolium. Nova Guinea neerlandica in reg. flum. Mamberamo, pr. voet Doormanstop. 3250m.(10663 ft.) http://data.biodiversitydata.nl/naturalis/specimen/L%20%200000475 And http://data.biodiversitydata.nl/naturalis/specimen/L%20%200000476 High epiphyte in mossy forest at 2150m. Branches erect, flowers greenish-white. 18 km SW. of Bernhard Camp, Taritatu (Idenburg) River, 1939. http://data.biodiversitydata.nl/naturalis/specimen/L.2913090 Doormantop, 1920. Type, http://data.biodiversitydata.nl/naturalis/specimen/L%20%200000477 Type. https://s3.amazonaws.com/huhwebimages/7700671B97FC45E/type/full/96797.jpg National History Museum. UK. http://data.nhm.ac.uk/dataset/collection-specimens/resource/05ff2255-c38a-40c9-b657-4ccb55ab2feb/record/467675 Queensland H. http://plants.jstor.org/stable/10.5555/al.ap.specimen.bri-aq0570114 Ecology/Infauna. A high-altitude species identified as LAM 1641 in Huxley (1978.) Large, often terrestrial, rarely contained ants but often frogs, lizards and large invertebrates. Huxley, C. R. “The Ant-plants Myrmecodia and Hydnophytum (Rubiaceae), and the relationships between their morphology, ant occupants, physiology and ecology.” Click PDF. http://onlinelibrary.wiley.com/doi/10.1111/j.1469-8137.1978.tb02285.x/citedby
  19. H. crassicaule P. Royen (Pieter van Royen) published in The Alpine Flora of New Guinea vol. 4, p2672/3, (1983) (Alpine Fl. New Guinea.) Bogor specimen relegated to H. alboviride http://bioportal.naturalis.nl/multimedia/L.2913091_01435743919/term=Hydnophytum+crassicaule&from=0 Royen Holotype relegated to H. alboviride. http://data.biodiversitydata.nl/naturalis/specimen/L.2913818 And. http://data.biodiversitydata.nl/naturalis/specimen/L.2913091 http://data.biodiversitydata.nl/naturalis/specimen/L%20%200000477
  20. H. vaccinifolium P. Royen. (Pieter van Royen) The alpine flora of New Guinea: volume 4. Taxonomic part, Casuarinaceae to Asteraceae, J. Cramer, (Alpine Fl. New Guinea 4, p2668, 1983.) Lam #1641 Originally labelled H. condensatum s. n., then in 1991. H. alboviride, and (undated) holotype of H. vaccinifolium. Nova Guinea neerlandica in reg. flum. Mamberamo, pr. voet Doormanstop. 3250 m. (10663 ft.) http://data.biodiversitydata.nl/naturalis/specimen/L%20%200000475 And http://data.biodiversitydata.nl/naturalis/specimen/L%20%200000476
  21. Here H. nigrescens is referred to Hydnophytum tortuosum by C. R. Huxley-Lambrick. Isotype Qld. H. http://plants.jstor.org/stable/10.5555/al.ap.specimen.bri-aq0570122
  22. It seems that H. auridemens may become a variety of H. petiolatum if the Jebb & Huxley revision is ever published. This is from Chomicki et al., support information. He obviously has access to the unpublished manuscript. I have added huge volumes of data to Epiphytic Myrmecophytes: Bizarre Wonders of Nature. 2019. Currently I am working on chapter 9 Hydnophytum. It alone contains over 88,000 kilobytes of data and images. H. auridemens Jebb & C. R. Huxley in press. Replaces manuscript name H. missima (sic.) H. petiolatum var. auridemens in press. Description. Tuber irregular, often containing rainwater. Bract hairs orange-brown, dark brown when wet. Habitat/Range. High-level epiphyte at 300-400 m (984-1312 ft.) in lower tropical forest to lower montane open forest, on south slopes of Mt Sisa, Misima Island, Louisiade Archipelago, Milne Bay Province, Papua New Guinea. Errata. One collection is mislabeled Hydnophytum aundemens Jebb 395, 1983 in http://www.pngplants.org/ database. Holotype. http://www.pngplants.org/specImages/LAE258387.jpg Isotype. http://apps.kew.org/herbcat/getImage.do?imageBarcode=K000761977 Johns et al. (2009). An introduction to the flora of the Milne Bay Archipelago. http://pharmacy.utah.edu/ICBG/pdf/WebResources/ForestBiodiversity/Johns-2009-Milne-Bay-flora.pdf
  23. Here are a few more records. Type http://kiki.huh.harvard.edu/databases/specimen_search.php?mode=details&id=5515 Huxley seems to have accepted it as a Hydnophytum? http://data.biodiversitydata.nl/naturalis/specimen/L%20%200000494 http://data.biodiversitydata.nl/naturalis/specimen/L%20%200000495 Bogor H. http://data.biodiversitydata.nl/naturalis/specimen/L.2913584
  24. Here is a clearer type specimen http://apps.kew.org/herbcat/getImage.do?imageBarcode=K000761965
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